You cannot enable the auto-responder, because you have set the option "Don't save incoming e-mails"

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If the check box "Don't save incoming e-mails" is selected in the mailbox edit form, the email server will ignore the incoming email messages, thus the auto-responder will not work.

To enable the auto-responder, clear the check box "Don't save incoming e-mails" in the mailbox edit form.

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